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That if you would take all the samples of that enzyme from a bunch of different organisms, in this case Saccharomyces cerevisiaeyeast and the water-related species, look at the various alcohol dehydrogenase enzymes, you could build, as he did-- his team did-- 12 different versions of the primordial ADH,what the first one looked like.
Listen to native speakers pronounce “dehydrogenase” as a noun in real conversational contexts with synchronized timestamps and subtitles.